Ch2oh

COO"

COO"

FIGURE 20-21 Glycolate pathway. This pathway, which salvages 2-phosphoglycolate (shaded pink) by its conversion to serine and eventually 3-phosphoglycerate, involves three cellular compartments. Glycolate formed by dephosphorylation of 2-phosphoglycolate in chloroplasts is oxidized to glyoxylate in peroxisomes and then transaminated to glycine. In mitochondria, two glycine molecules condense to form serine and the CO2 released during photorespiration (shaded green). This reaction is catalyzed by glycine decarboxylase, an enzyme present at very high levels in the mitochondria of C3 plants (see text). The serine is converted to hydroxypyruvate and then to glyc-erate in peroxisomes; glycerate reenters the chloroplasts to be phos-phorylated, rejoining the Calvin cycle. Oxygen (shaded blue) is consumed at two steps during photorespiration.

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