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propionyl CoA; and (2) simple a-keto acids such as pyruvate. Each substrate must contain a carbonyl group adjacent to or conjugated with the carbon receiving the carboxyl group from carboxy biocytin. Perhaps the most familiar biotin carboxylase enzymes in mammalian systems are acetyl-CoA carboxylase in fatty acid biosynthesis, propionyl CoA carboxylase in odd-chain fatty acid catabolism, pyruvate carboxylase in gluconeogenesis, and ^-methyl-crotonyl CoA carboxylase in leucine catabolism.

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